Set in the fictional Florida town of Mangrove Bay during the 1940s, the novel follows a Black family and their close friends as they navigate racial segregation, military service in World War II, and the broken promises of postwar America.
Benny North, a light-skinned Black man who can pass for white, uses this ability to access segregated gas stations and other whites-only spaces in the Jim Crow South. He lives with his sister Cora and their mother, Momma (Janie) North, a hotel maid. The family survived the 1923 Rosewood massacre, a real act of racial violence in Florida that killed their father. Benny's closest friends include Roscoe Crane, his lifelong best friend; Jasper Monroe, a trumpet-playing parks attendant; and Lee Peters, a gifted musician and Negro League catcher whose parents were murdered for their political activism. Lee's defense lawyer, Uncle Drew (Drew Brooks), became his guardian.
Cora and Lee carry on a secret romance, hiding it from Momma, their Aunt Teen, and their church community, who consider Lee unworthy because of his juvenile record. On December 7, 1941, the attack on Pearl Harbor upends everything. Benny, Roscoe, and Lee talk of enlisting, while Momma, Cora, and Jasper argue that Black men owe nothing to a country that has never treated them as citizens. Lee tells Cora he is enlisting, believing military service will earn him the respect needed for them to be together openly. She begs him to stay, but he leaves without asking her to wait, a silence that wounds her deeply.
With Lee gone, Roscoe proposes a plan: He will marry Cora so that if he or Benny dies in the war, she will receive military widow's benefits. Roscoe has been attracted to Cora for years. Cora, heartbroken and doubting Lee's commitment, accepts out of duty and love for Benny. The February 1942 courthouse wedding is quick and loveless, and the marriage remains unconsummated.
The three men's military experiences diverge sharply. Benny is mistakenly placed in an all-white camp. His commanding officer, Captain Bale, discovers the discrepancy but crosses out "colored" and writes "white," ordering Benny to keep silent. Benny bonds with his white comrades through combat in North Africa, terrified of exposure but accepted as one of their own. Roscoe is assigned to menial labor before transferring to the 320th Barrage Balloon Battalion, a unit training Black soldiers to operate hydrogen-filled balloons designed to block enemy bombers. Lee joins the 761st Tank Battalion, the Black Panthers, where racist officers subject the men to abuse. Lieutenant Colonel Paul Bates transforms the unit, refusing a promotion to stay with his men and defending Second Lieutenant Jackie Robinson after Robinson is arrested for refusing to give up his bus seat.
Back home, Cora works at Sunshine State Insurance alongside another Black employee in a windowless closet while white secretaries enjoy a spacious office. Her cousin Patsy, a trained nurse, is rejected by the Army Nurse Corps because the army caps Black nurses at fifty-six. Cora launches a letter-writing campaign, and the quota is eventually raised. When Cora visits Patsy's posting, she discovers Patsy has been assigned to care for German prisoners of war who enjoy freedoms denied to Black Americans.
Unable to confess her marriage, Cora avoids writing to Lee for over a year. When she finally sends a letter revealing the truth, Lee is devastated. He channels his heartbreak into combat after the 761st deploys to Europe. At the Battle of the Bulge, Benny and Lee encounter each other. Benny calls out but freezes, remembering he is supposed to be white. They exchange a stiff handshake instead of their old greeting and part like strangers. Jasper, fighting in Italy with the Buffalo Soldiers 92nd Division, is killed in action. After Germany surrenders, Lee and the Black Panthers liberate the Gunskirchen concentration camp. When Lee overhears a white officer say America needs concentration camps for Black people, he punches the man unconscious. The army issues Lee a blue discharge, neither honorable nor dishonorable, stripping him of all GI benefits.
Roscoe, deployed to Wales, experiences life without segregation for the first time and falls in love with a local woman named Megan Davies. After the war, Lee returns to Mangrove Bay, reconnects with Cora, and throws himself into building Green's Whiskey, a distillery based on the family recipe of his fallen comrade Green George. Roscoe arrives home to find Lee established in Cora's household and turns to heavy drinking, steered by the Veterans Affairs office into degrading farm labor.
Benny settles into Levittown, a whites-only planned community, taking full advantage of GI Bill benefits denied to Black veterans. He falls in love with his neighbor's sister, Gloria, who denounces segregation as "just plain wrong" (256). Cora, fired from Sunshine State because returning white workers make her presence unacceptable, joins Lee at Green's Whiskey and launches an advocacy campaign with Uncle Drew, speaking to the press about housing discrimination. When Cora discovers four crumpled, unfinished letters from Roscoe addressed to Megan, she confronts him. Drunk and furious, Roscoe strikes her across the face. Cora refuses to cower, goes to Lee, and they finally begin their life together.
White vigilantes attack Green's Whiskey twice. The second time, five men overpower Lee, tie him to their truck bumper, and drag him through the streets. Cora escapes through a window and runs after the truck on glass-shredded feet. The whiskey barrels ignite and the building explodes. Benny, driving through a hurricane, finds Cora collapsed on the roadside, locates the attackers' truck in a ditch, and cuts Lee free. Lee's injuries are catastrophic, including dislocated shoulders, a shattered hip, and skin scraped to bone. He falls into a coma for weeks.
With Green's Whiskey destroyed and Roscoe gone, the family faces eviction. Cora convinces Benny to use his white-passing status to obtain a GI business loan and buy the burned-out property to build Liberty Heights, a Black housing development. Uncle Drew finds a loophole: The Federal Housing Administration (FHA) loan requires selling to a "harmonious racial group," which an all-Black community satisfies. The FHA investigates when Black buyers appear, but Uncle Drew argues that Benny followed the contract to the letter and that recent Supreme Court rulings invalidate racial restrictions. The agency drops the case but bars Benny from future government loans.
Lee wakes from his coma. Doctors tell him he may never walk again, and he tries to push Cora away, but she refuses. She brings him to a house at Liberty Heights designed for wheelchair accessibility, with ramps, lowered counters, and handrails. Roscoe sails back to Wales, sending Cora divorce papers. Benny, unable to sustain his double life, breaks up with Gloria and sells his Levittown house. Before reenlisting in the newly desegregated military under President Truman's 1948 executive order, he returns to confess his identity to Gloria. She is shocked and angry about the lies but tells him he can write to her and promises she will answer.
In an epilogue set on June 25, 1950, the community gathers at Lee and Cora's Liberty Heights home for a cookout. Patsy is pregnant, married to Dr. Allen George, the physician who treated Lee. Lee can now stand and take small steps after two years of therapy. Cora learns Roscoe has had a son in Wales named Jasper, after their fallen friend. Gloria arrives carrying a Key lime pie, stepping into Benny's family's home for the first time. Lee plays Jasper's old trumpet under the orange tree he planted for Cora, filling the yard with music that carries the weight of everything they have survived.